While Frostpaw takes refuge in ShadowClan as Splashtail seizes control of her home, Nightheart returns to ThunderClan and to a complicated reunion with his mate Sunbeam whose efforts to fit in as ThunderClan's newest member are hampered by her treacherous mother.

"In the Clandoms, everyone is born with wings, though not all the clans are treated equally. Ellie Meadows longed to become a Goldwing--the knights who protect the people--but because she came from the humble Sparrow Clan, her dream faced almost insurmountable resistance. Nox Hatcher has it even worse. Nox is a Crow, a shattered clan. They can't own businesses or run organizations, all due to a mysterious betrayal made hundreds of years ago. Nox has...

Raised to be a warrior, seventeen-year-old Eelyn fights alongside her Aska clansmen in an ancient rivalry against the Riki clan, but when faced with her brother's betrayal, and driven by a growing love for her brother's friend Fiske, she attempts to unite the two clans.

When an outsider appears in the ThunderClan's midst, it drives the Clans further apart and as tension mounts they must decide who they can trust before it is too late.